Dianthus gracilis Sm. subsp. gracilis

Taxonomy: Fam. Caryophyllaceae// Genus: Dianthus// Species: Dianthus gracilis Sm.// Subspecies: Dianthus gracilis Sm. subsp. gracilis.

Determined according to the botanical key provided by (Micevski, 1993) and Tutin in Tutin et al. (1964).

Two subspecies of Dianthus gracilis Sm. occur in the Osogovo region: Dianthus gracilis Sm. subsp. gracilis and Dianthus gracilis subsp. armerioides (Griseb.) Tutin. Distinction between subspecies:

Dianthus gracilis Sm. subsp. gracilisEpicalyx-scales reaching up to c. 1/2 of the calyx length. Bracts very short, reaching the lower 1/3 of the calyx;

Dianthus gracilis subsp. armerioides (Griseb.) Tutin: Epicalyx-scales reaching min 3/4 of the calyx length. Bracts conspicuously longer- as long as the calyx or almost so.

Lifeform: Perennial, sub-shrub; chamaephyte according to the Raunkiaer system.

Habitat: Found from the lowland up to mid-altitudes (c. 800-900 m) mainly on rocky substrates in open and semi-open habitats- dry grasslands, edges of forests, rocky terrains, river gorges.

Distribution: Native to the Balkan peninsula according to The Euro+Med PlantBase (Marhold, 2011+) and the Plants of the World Online portal (Royal Botanic Gardens, Kew).

Flowering period: Summer. In favorable condition plans can be found in bloom from late June until October. Peak flowering occurs in July: JUN, JUL, AUG, SEP (-OCT).

Dianthus gracilis subsp. gracilis – habit. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – habit. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – habit. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – habit. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – habit. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flowering stem with flowers. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flowering stem with flowers. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flowering stems with flowers. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flowering stem with spent flowers. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flowering stem with flowers. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flower. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flower. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flower. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flower. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flower. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flower. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flower – calyx, epicalyx and bracts. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flowers – calyx, epicalyx and bracts. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flowers – calyx, epicalyx and bracts. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – flowers – calyx, epicalyx and bracts. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – calyx, epicalyx and bracts. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – calyx and epicalyx. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – stem and cauline leaves . Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – leaf sheath of cauline leaves. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – abaxial side of leaf. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – lower part of flowering stems and vegetative stems. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – vegetative stems. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – vegetative stems. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – leaves of vegetative stems. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.
Dianthus gracilis subsp. gracilis – plant base- woody stems. Gorge of “Kochanska” River, 16.07.2022. Photo by Trajche Dimitrovski.

References:

Dianthus gracilis subsp. gracilis. Marhold, K. (2011+): Caryophyllaceae. – In: Euro+Med Plantbase – the information resource for Euro-Mediterranean plant diversity. Published on the Internet. https://europlusmed.org/cdm_dataportal/taxon/d41f88df-3180-48fc-8ce8-903380fe36d9 [Accessed 22.01.2023].

Dianthus gracilis subsp. gracilis. Plants of the World Online, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:77229077-1 [Accessed 22.01.2023].

Dianthus L. Micevski, K. (1993). The Flora of the Republic of Macedonia Vol. I, Book 2. Macedonian Academy of Sciences and Arts, pp 360 – 390.

Dianthus L. Tutin, T.G. in Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M., & Webb, D.A. (1964). Flora Europaea. Volume 1. Lycopodiaceae to Platanaceae. Cambridge: Cambridge University Press, pp. 188-204.

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