Determined according to the botanical key by Fernandes (1976 in Tutin et al.) and the distinguishing traits for the Anthemis macedonica Group provided by Goula at al. (2022). According to the taxonomic concept accepted by The Euro+Med PlantBase (Greuter, 2006+), Anthemis macedonica Boiss. & Orph. is treated as a polymorphic species with 4 subspecies. According to the current concept, the populations on Osogovo belong to the autonym subspecies.
Lifeform: Annual (therophyte according to the Raunkiær system) or biennial plant.
Distribution: Native to the Balkan Peninsula according to The Euro+Med PlantBase (Greuter, 2006+) and the Plants of the World Online portal (Royal Botanic Gardens, Kew).
Habitat: Open and dry habitats – hill pastures, dry grassland, rocks and rocky terrains with shallow soil, river gorges.
Flowering period: Spring to summer (MAY, JUN, JUL).
Anthemis macedonica – habit. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – capitula. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – capitula. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – involucral bracts. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – stems with leaves. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – stem and upper cauline leaves. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – leaves. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – leaves. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – lower leaf. Gorge of “Kochanska” River, 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica. Scale in cm. Specimen from the gorge of “Kochanska” River, collected 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica. Scale in cm. Specimen from the gorge of “Kochanska” River, collected 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica. Scale in cm. Specimen from the gorge of “Kochanska” River, collected 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica. Scale in cm. Specimen from the gorge of “Kochanska” River, collected 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum (left) and involucral bracts and underside of petals (right). Scale in cm/mm. Specimen from the gorge of “Kochanska” River, collected 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – upper, middle and lower leaf (from left to right). Scale in cm/mm. Specimen from the gorge of “Kochanska” River, collected 24.05.2025. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – upper part of plant with capitula. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum with involucre. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – stem with cauline leaves. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – stem with cauline leves. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – cauline leaf. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – plant base. Gorge of “Kochanska” River, 22.05.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 20.06.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – habit. Gorge of “Kochanska” River, 20.06.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – plant senescence during achene ripening. Gorge of “Kochanska” River, 20.06.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum with developing achenes. Gorge of “Kochanska” River, 20.06.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum with developing achenes. Gorge of “Kochanska” River, 20.06.2021. Photo by Trajche Dimitrovski. Anthemis macedonica – capitulum with developing achenes. Gorge of “Kochanska” River, 20.06.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum with mature achenes. Gorge of “Kochanska” River, 20.06.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – fruiting capitulum. Gorge of “Kochanska” River, 20.06.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – plant senescence during achene ripening. Gorge of “Kochanska” River, 29.08.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – capitulum with scales. Gorge of “Kochanska” River, 28.09.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – achene. Specimen from the Gorge of “Kochanska” River, accessed 28.09.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – achene. Specimen from the Gorge of “Kochanska” River, accessed 28.09.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – achenes. Specimen from the Gorge of “Kochanska” River, accessed 28.09.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – achene.Specimen from the Gorge of “Kochanska” River, accessed 28.09.2021. Photo by Trajche Dimitrovski.Anthemis macedonica – achene. Specimen from the Gorge of “Kochanska” River, accessed 28.09.2021. Photo by Trajche Dimitrovski.
Fernandes, R. 1976. Anthemis L. In Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A. (eds.), Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae). Cambridge: Cambridge University Press, pp. 145–159.
Goula, K., Touloumis, K., Dimopoulos, P. & Constantinidis, T. A Morphometric and Karyological Study of the Anthemis macedonica Group (Asteraceae, Anthemideae) Reveals a New Species from Greece. Plants, 11(21), 3006. https://doi.org/10.3390/plants11213006 [Accessed 19.01.2026].