Xanthium orientale subsp. italicum (Moretti) Greuter (syn. Xanthium strumarium subsp. italicum (Moretti) D. Löve)

Taxonomy: Fam. Compositae// Genus:  Xanthium// Species: Xanthium orientale L.// Subspecies: Xanthium orientale subsp. italicum (Moretti) Greuter

Determined according to the botanical key provided in Flora Europaea Vol. 4 (Tutin et al., 1976) as Xanthium strumarium subsp. italicum (Moretti) D. Löve. According to The Euro+Med PlantBase (source: Greuter, 2006+) this taxon is a synonym of Xanthium orientale subsp. italicum (Moretti) Greuter.

Lifeform: Annual; therophyte according to the Raunkiær system.

Distribution: Native to South and North America, introduced in Europe (Tutin et al., 1976). Naturalized alien in North Macedonia according to The Euro+Med PlantBase (source: Greuter, 2006+).

Habitat: In the Osogovo region this species is a usually associated with disturbed ground; generally found in the ruderal area of the region. Common on waste places, roadsides, cultivated land, corn plantations.

Flowering period: Summer – autumn (JUL, AUG, SEP).

Xanthium orientale subsp. italicum –stem with fruits and leaves. Roadside near “Kochanski dol” locality, Kochani, 19.09.2021. Photo by Trajche Dimitrovski.
Xanthium orientale subsp. italicum –stem with fruits . Roadside near “Kochanski dol” locality, Kochani, 19.09.2021. Photo by Trajche Dimitrovski.
Xanthium orientale subsp. italicum –stem with fruits . Roadside near “Kochanski dol” locality, Kochani, 19.09.2021. Photo by Trajche Dimitrovski.
Xanthium orientale subsp. italicum – fruits . Roadside near “Kochanski dol” locality, Kochani, 19.09.2021. Photo by Trajche Dimitrovski.
Xanthium orientale subsp. italicum – fruits . Roadside near “Kochanski dol” locality, Kochani, 19.09.2021. Photo by Trajche Dimitrovski.
Xanthium orientale subsp. italicum – leaf . Roadside near “Kochanski dol” locality, Kochani, 19.09.2021. Photo by Trajche Dimitrovski.
Xanthium orientale subsp. italicum – leaf . Roadside near “Kochanski dol” locality, Kochani, 19.09.2021. Photo by Trajche Dimitrovski.

References:

Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (1976). Flora Europaea. Volume 4: Plantaginaceae to Compositae (and Rubiaceae). Cambridge: Cambridge University Press.

Xanthium orientale subsp. italicum. Greuter, W. (2006+): Compositae (pro parte majore). – In: Greuter, W. & Raab-Straube, E. von (ed.): Compositae. Euro+Med Plantbase – the information resource for Euro-Mediterranean plant diversity. Published on the Internet: http://ww2.bgbm.org/EuroPlusMed/PTaxonDetail.asp?NameId=7530648&PTRefFk=7000000 [Accessed 17.10.2021].

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