Myosotis sylvatica subsp. cyanea (Hayek) Vestergr.

Posted byTrajche DimitrovskiPosted inBoraginaceae, Plant families

Taxonomy: Fam. Boraginaceae// Genus: Myosotis// Species: Myosotis sylvatica Hoffm.// Subspecies:  Myosotis sylvatica subsp. cyanea (Hayek) Vestergr.

Determined according to the botanical key provided by Matevski and Melovski (2010 in Matevski).

Lifeform: Biennial plant. First year growth (vegetative stage) is characterized by basal rosette formation. Flowering occurs in the second year.

Distribution: Native from Southeast Europe to Transcaucasus according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Gardens, Kew).

Habitat: Grasslands and pastures, clearings and edges of forests, along streams and rivers and other wet habitats in the zones of European beech forests and subalpine pastures. Common member of the hydrophilous (tall herb fringe) communities along streams and rivers in the upper part of Osogovo Mountain. Recorded from c. 750 up to 1950 m altitude on several localities: “Shtalkovichka” River, Ponikva, “Carev vrv” peak region.

Flowering period: Spring to early summer (APR, MAY, JUN).

Locality: Ponikva.

Myosotis sylvatica subsp. cyanea – habit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – habit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – habit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – habit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – habit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – upper part with inflorescence. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowering stem with inflorescence. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – inflorescence. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyx. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyces in fruit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyx in fruit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyx in fruit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyces in fruit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyces in fruit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – axis of inflorescence with appressed hairs; calyces in fruit. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – axis of inflorescence with appressed hairs. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stems with cauline leaves. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stems with cauline leaves. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem with cauline leaves. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem with cauline leaves. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – cauline leaf. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea– adaxial side of leaf. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – adaxial side of leaf. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – abaxial side of leaf. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea– abaxial side of leaf. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – basal leaves and stem with cauline leaves. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – lower part of stem with basal and lower cauline leaves. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – basal leaves. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – basal leaf. Osogovo, Ponikva, 28.05.2022. Photo by Trajche Dimitrovski.

Scaled images (specimens collected in Ponikva, 08.06.2025).

Myosotis sylvatica subsp. cyanea – habit. Scale in cm. Specimen from Osogovo, Ponikva. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers and fruiting calyces. Scale in mm. Specimen from Osogovo, Ponikva. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers and fruiting calyces. Scale in mm. Specimen from Osogovo, Ponikva. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stems with leaves. Scale in mm. Specimen from Osogovo, Ponikva. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – hooked calyx hairs. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – hooked calyx hairs. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – hooked calyx hairs. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – hooked calyx hairs. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – hairs of calyx tooth. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – hairs of pedicel. Photo by Trajche Dimitrovski.

Locality: “Shtalkovichka” River (Mangov kamen).

Myosotis sylvatica subsp. cyanea – habit. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – inflorescence. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – inflorescence. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – fowers. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – fowers. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – fowers. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – fowers. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyces and axis of inflorescence. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyces and axis of inflorescence. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem and leaf. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – fruiting calyces. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem with leaves. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem with leaves. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – lower leaf. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – abaxial side of leaf. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem with leaves. Osogovo – “Shtalkovichka” River, 07.05.2023. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyx hairs under magnification. Specimen from Osogovo – “Shtalkovichka” River. Photo by Trajche Dimitrovski.

Locality: “Carev vrv” peak.

Myosotis sylvatica subsp. cyanea – habit. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – habit. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowering stems. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – flowers. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyces. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – calyces. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – fruiting calyces and axis of inflorescence. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem and leaf. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem with leaves. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem with leaves. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – stem and adaxial side of leaf. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp. cyanea – abaxial side of leaf. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.
Myosotis sylvatica subsp.cyanea – lower part of plant. Osogovo – “Carev vrv” peak, 09.06.2024. Photo by Trajche Dimitrovski.

References:

Matevski, V. and Melovski, Lj. 2010. In Matevski, V. (ed.), The Flora of the Republic of Macedonia. Vol. II, Book 1. Skopje: Macedonian Academy of Sciences and Arts, pp. 141–163.

Myosotis sylvatica subsp. cyanea (Hayek) Vestergr. Plants of the World Online, Royal Botanic Gardens Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:77252061-1 [Accessed 29.05.2022].

Myosotis sylvatica subsp. cyanea. Valdés, B. & Raab-Straube, E. von 2011+: Boraginaceae. – In: Euro+Med PlantBase – the information resource for Euro-Mediterranean plant diversity. Published at https://europlusmed.org/. https://europlusmed.org/cdm_dataportal/taxon/e0e6418b-ca0b-40ea-ab86-bac497f0069e [Accessed 14.01.2026].

Last update: 14.01.2026.