Pisum sativum subsp. elatius (M. Bieb.) Asch. & Graebn.

Taxonomy: Fam. Fabaceae// Genus: Pisum// Species: Pisum sativum L.// Subspecies: Pisum sativum subsp. elatius (M. Bieb.) Asch. & Graebn.

Determined according to the botanical key provided by Micevski in Micevski (2001).

Lifeform: Annual; therophyte according to the Raunkiær system.

Distribution: According to Zlatkovic, Mikic and Smykal (2010), Pisum sativum L. subsp. elatius is a widely dispersed Mediterranean element with native range spreading across the Mediterranean and Oriental-Thuranian floristic regions. In the Plants of the World Online portal (Royal Botanic Gardens, Kew), both subspecies of Pisum sativum L. are treated as synonyms of Lathyrus oleraceus Lam. with native distribution from Mediterranean to Afghanistan.

Habitat: Clearings of oak forests, rock cliffs at c. up to c. 900 – 950 m altitude.

Flowering period: Late spring – early summer: MAY, JUN.

*Pisum sativum* subsp. *elatius* – habit. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – stems with inflorescences. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – stem with inflorescences. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – flower. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – calyx. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – developing flower. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – lyx. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – plant with developing pods. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – pods. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – pod. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – immature seeds. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – stem with leaves and stipules. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – adaxial side of leaf. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – abaxial side of leaf. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – tendril. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – stipules. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – stem. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – plant base. Osogovo – “Kochanski dol” near village of Shtalkovica, 01.06.2025. Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – left: pod and leaf with stipules (scale in cm); right: width of pods (scale in mm). Photo by Trajche Dimitrovski.

*Pisum sativum* subsp. *elatius* – immature seeds (A) and mature seed (B). Scale in mm; surface of seed (C). Photo by Trajche Dimitrovski.

References:

Lathyrus oleraceus Lam. Plants of the World Online portal, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:501912-1 [Accessed 21.06.2025].

Pisum L. Micevski, K. (2001). The Flora of the Republic of Macedonia Vol. I, Book 5. Macedonian Academy of Sciences and Arts, pp. 1255 – 1256.

Zlatkovic, B., Mikic, A. and Smykal, P. (2010). Distribution and new records of Pisum sativum subsp. elatius in Serbia. Pisum genetics, Vol. 42, pp. 15 – 17. Dowloaded from https://www.cabidigitallibrary.org/doi/pdf/10.5555/20123231291 [Accessed 21.06.2025].

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