Determined according to the botanical key provided in Flora Europaea (Tutin et al., 1976).
Lifeform: Perennial; Hemicryptophyte according to the Raunkiær system.
Distribution: Native from Central Europe to Italy, Balkan Peninsula and Romania according to the Euro+Med PlantBase (Greuter, 2006+).
Habitat: found in somewhat shaded habitats – mixed oak forests, near rivers.
Flowering period: Spring: APR, MAY.
Doronicum columnae– habit in habitat (Querco-carpinetum orientalis forest community). Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit in habitat (Querco-carpinetum orientalis forest community). Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– capitulum. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– capitulum. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– capitulum – florets. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– involucre of capitulum. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– stem with cauline leaves and basal leaves. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– cauline leaf. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– basal leaf. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– basal leaves. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– rootstock. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– rootstock – tubers with sericeous hairs. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– rootstock – tubers with sericeous hairs. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– capitulum. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– capitulum. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– involucre of capitulum. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– capitulum with developing achenes. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– cauline leaves. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– cauline leaves. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– stem with cauline leaves and basal leaves. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– basal leaves. Osogovo, moutain ridge between lake “Gradche” and the gorge of “Orizarska” River, 08.05.2022. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.Doronicum columnae– habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.Doronicum columnae– capitulum. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.Doronicum columnae– capitulum. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski. Doronicum columnae– capitulum. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.Doronicum columnae– involucral bracts. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.Doronicum columnae– involucral bracts of developing capitulum capitulum. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski. Doronicum columnae– stem with cauline leaves and basal leaves. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski. Doronicum columnae– cauline leaf. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski. Doronicum columnae– cauline leaf. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski. Doronicum columnae– cauline leaf and basal leaves. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski. Doronicum columnae– basal leaves. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.Doronicum columnae– basal leaves. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski. Doronicum columnae– rhizome with tufts of sericeous hairs. Gorge of “Kochanska” River, 29.04.2021. Photo by Trajche Dimitrovski. Doronicum columnae– rhizome with tufts of sericeous hairs. Gorge of “Kochanska” River, 29.04.2021. Photo by Trajche Dimitrovski. Doronicum columnae– rhizome with tufts of sericeous hairs. Gorge of “Kochanska” River, 29.04.2021. Photo by Trajche Dimitrovski.
Osogovo nature 