Determined according to the botanical key provided in Werner in Tutin et al. (1976) to species level and Gajić in Josifović (1975) to subspecies level.
Species variability: Two subspecies of Cirsium ligulareBoiss. occur in the Osogovo region: Cirsium ligulare subsp. albanum Wettst. and Cirsium ligulare subsp. armatum (Velen.) Petr. Distinction between subspecies:
–Cirsium ligularesubsp. albanumWettst:Involucral bracts distinctly widened (lanceolate) below the apical spine; apical spine up to 6 mm long. Widely distributed from the lowland up to 1600 m;
– Cirsium ligularesubsp.armatum(Velen.) Petr.: Involucral bracts gradually narrowed into the apical spine, not distinctly widened in the apical part; the apical spine usually longer as compared to subsp. albanum; found in the higher parts of the Osogovo region;
Lifeform: Biennial; the first year of the lifecycle is vegetative (basal rosette stage). Flowering and fruiting occurs in the second year.
Distribution: Native from the Balkan Peninsula according to the distribution information provided by The Euro+Med PlantBase (source: Greuter, 2006+) and the Plants of the World Online portal (Royal Botanic Gardens, Kew).
Habitat: Grasslands and pastures, from the lowland up to c. 1600 m altitude.
Flowering period: Summer- autumn: AUG, SEP, OCT.
Cirsium ligulare subsp. albanum – habit. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – flowering stems with capitula. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – capitulum. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – capitulum. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – florets. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – involucral bracts. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – stem with cauline leaves. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – cauline leaves. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – cauline leaf. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – adaxial side of leaf. Osogovo – Ponikva, 15.08.2022. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– habit. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– habit. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– upper part of plant with capitula. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– capitula. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– developing capitulum. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– developing capitulum. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– capitulum. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– capitulum. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– capitulum. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – capitulum. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – spent capitulum. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum – spent capitulum (left) and fruiting capitulum (right). Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– capitulum with achenes. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– cauline leaf. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– stem with cauline leaves. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– stem with cauline leaves. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligularesubsp. albanum– stem with cauline leaves. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– habit of young plant in vegetative stage- first year growth. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.Cirsium ligulare subsp. albanum– basal leaf. Osogovo, hilly region near village of Beli, 20.09.2020. Photo by Trajche Dimitrovski.
References:
Cirsium ligulare subsp. albanum. Greuter, W. (2006+): Compositae (pro parte majore). – In: Greuter, W. & Raab-Straube, E. von (ed.): Compositae. Euro+Med Plantbase – the information resource for Euro-Mediterranean plant diversity. https://europlusmed.org/cdm_dataportal/taxon/7aebd00a-83ca-4535-b94d-2dfbf436f737 [Accessed 26.01.2023].