Determined according to the botanical key provided by Micevski (1985), Vukićević in Josifović, 1970 and Crabbe et al. in Tutin et al. (1964) as Asplenium cuneifolium Viv. The accepted name in the Euro+MedPlantBase is Asplenium adiantum-nigrum subsp. serpentini (Tausch) Heufl.
Lifeform: Perrenial; Hemicryptophyte according to the Raunkiær system.
Distribution: Native from Europe to Iran according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Garden, Kew).
Habitat: Shaded positions on rocks- Shtalkovicka River and “Ratkova Skala” region.
Asplenium adiantum-nigrumsubsp. serpentini – habit. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – habit. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – habit – dried fronds from the pervious year and new developing fronds. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – habit. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – leaf blades. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – leaf blades. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – terminal part of blade – rachis and pinnae with pinnules. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – rachis and pinnae with pinnules. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – rachis and pinnae with pinnules. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini –rachis and pinna with pinnules. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – pinna with pinnules. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – abaxial side of pinna with developing sori. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – abaxial side of pinna with developing sori. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – pinna with sori of leaf from the previous year. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrumsubsp. serpentini – petioles. Osogovo – “Ratkova Skala” region between the the villages of Shtalkovica and Ratkovica, 29.04.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. serpentini – habit. “Shtalkovichka” River, 27.03.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. serpentini – leaf. “Shtalkovichka” River, 27.03.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. serpentini – pinnae. “Shtalkovichka” River, 27.03.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. serpentini – pinnae. “Shtalkovichka” River, 27.03.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. serpentini – pinnae. “Shtalkovichka” River, 27.03.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. serpentini – abaxial side of pinnae with sori. “Shtalkovichka” River, 27.03.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. serpentini – abaxial side of pinnae with sori. “Shtalkovichka” River, 27.03.2023. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. serpentini – leaf petiole. “Shtalkovichka” River, 27.03.2023. Photo by Trajche Dimitrovski.
Osogovo nature 