Taxonomy: Fam. Pteridaceae// Genus: Anogramma// Species: Anogramma leptophylla (L.) Link. The classification is given according to The Euro+Med PlantBase (source: Christenhusz & Raab-Straube, 2013+).
Determined according to the botanical keys provided by Micevski (1985), Valentine (1964 in Tutin et al., Gymnogrammaceae) and Tutin (1964 in Tutin et al., Anogramma).
Lifeform: Annual fern. Therophyte according to the Raunkiær system. Note: sporophyte annual, gametophyte perennial according to Tutin (1964 in Tutin et al.).
Distribution: Native from Central Europe to Subtropics and Tropics – Asia, Africa, Australia, Mexico to parts of South America according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Garden, Kew).
Habitat: Moist positions on rock cliffs and cervices above Shtalkovichka River (“Ratkova Skala” region).
Locality: Osogovo , rock cliffs between the village of Ratkovica and the Shtalkovichka River. Date: 07.06.2026.
Habitat of Anogramma leptophylla. Osogovo , rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Habitat of Anogramma leptophylla. Osogovo , rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – leaves. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – adaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – adaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – adaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – adaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – adaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – adaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – adaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – dimorphic leaves. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – dimorphic leaves. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – dimorphic leaves. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.Anogramma leptophylla – abaxial side of leaves with sori. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 07.06.2026. Photo by Trajche Dimitrovski.
Locality: Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River. Date: 29.04.2023.
Habitat of Anogramma leptophylla. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo,ock cl riffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – plant with outer shorter leaves. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – inner longer leaves. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – inner longer leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – lamina of inner longer leaf. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – adaxial side of leaf with sori and sori under magnification. Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.Anogramma leptophylla – habit of plant with developed dimorphic leaves. Scale in mm. Specimen from Osogovo, rock cliffs between the village of Ratkovica and the Shtalkovichka River, 29.04.2023. Photo by Trajche Dimitrovski.
Micevski, K. (1985). Anogramma Link, The Flora of SR Macedonia. Vol. 1, Book 1. Skopje: Macedonian Academy of Sciences and Arts, pp. 52.
Tutin, T. G. 1964. Anogramma Link. In Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A. (eds.). Flora Europaea, Volume 1. Lycopodiaceae to Platanaceae. Cambridge: Cambridge University Press, p. 11.
Valentine, D. H. 1964. Gymnogrammaceae. In Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A. (eds.). Flora Europaea. Volume 1. Lycopodiaceae to Platanaceae. Cambridge: Cambridge University Press, p. 11.