Taxonomy: Fam. Caryophyllaceae// Genus: Cerastium// Species: Cerastium arvense L. // Subspecies: Cerastium arvense L. subsp. arvense.
Determined according to the botanical keys provided by Jalas; Sell and Whitehead (1964 in Tutin et al.) and Gajić (1970 in Josifović).
Habitat: Open habitats – subalpine pastures at c. 1600 – 1900 m altitude – Ponikva, Retki Buki, Mrtvica, the slopes below “Carev vrv” peak – Markova Stupka, Babina cheshma.
Distribution: Native from Europe to Central Asia, Russian Far East and Japan; Northwest Africa according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Gardens, Kew).
Lifeform: Perennial, hemicryptophyte according to the Raunkiaer system.
Flowering period: Late spring- summer: MAY, JUN, JUL.
Locality: Retki buki, 09.06.2024.
Cerastium arvensesubsp.arvense – habit in habitat. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit in habitat. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit . Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering stems arising from creeping stem. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering stems arising from creeping stem. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – inflorescence. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flower. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flower. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowers. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowers. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowers. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – side view of flowers with calyx; pedicels and bracts. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvense subsp. arvense – side view of flowers with calyx; pedicels and bracts. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – side view of flowers with calyx; pedicels and bracts. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – inflorescence. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – calyx with developing capsule. Fruiting pedicel curved just beneath the calyx. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – calyx with developing capsule. Fruiting pedicel curved just beneath the calyx. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – calyx with developing capsule. Fruiting pedicel curved just beneath the calyx. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering and non-flowering stems. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering and non-flowering stems. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering stems with leaves. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering stem with leaves. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering stem with leaves. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – abaxial side of leaf. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering stem with non-flowering axillary shoots. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves of non-flowering shoot. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves of non-flowering shoot. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – plant base – procumbent part of stem with flowering and vegetative shoots arising from it. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – procumbent part of stem. Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.
Cerastium arvensesubsp.arvense – habit. Scale in cm. Specimen collected from Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Scale in cm. Specimen collected from Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Scale in cm. Specimen collected from Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Scale in cm. Specimen collected from Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Scale in cm. Specimen collected from Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – fully developed flower (left), side view of developing flower with calyx (center) and capsule (right). Scale in mm. Specimen collected from Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering stem with cauline leaves (left), lower part of flowering stem with sterile shoots (center) and vegetative shoot (right). Scale in mm. Specimen collected from Osogovo – “Retki buki”, 09.06.2024. Photo by Trajche Dimitrovski.
Locality: Markova stupka, 15.07.2023.
Cerastium arvensesubsp.arvense – habit in habitat. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – habit. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering stems with inflorescence. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowering stems with inflorescence. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – inflorescence. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowers. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowers. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowers. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – flowers. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – side view of flowers and calyx; bracts. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – side view of flowers and calyx; bracts. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – side view of flowers and calyx; bracts. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – capsules enclosed by the calyx and bracts. The capsule is equal to the calyx length. The pedicels are curved beneath the calyx. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – capsule enclosed by the calyx and bracts. The capsule is equal to the calyx length. The pedicels are curved beneath the calyx. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – capsules enclosed by the calyx and bracts. The capsule is equal to the calyx length. The pedicels are curved beneath the calyx. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – capsule enclosed by the calyx and bracts. The capsule is equal to the calyx length. The pedicels are curved beneath the calyx. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – upper part of stem and bracts. The capsule is equal to the calyx length. The pedicels are curved beneath the calyx. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – vegetative stems. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – vegetative stems. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – abaxial side of leaves. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – abaxial side of leaves. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.
Scaled images (specimens from Markova stupka).
Cerastium arvensesubsp.arvense – capsules. The capsules are curved. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – capsules. The capsules are curved. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves of flowering stem. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves of flowering stem and non-flowering axillary shoots. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves of flowering stem. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves of non-flowering axillary shoots. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – vegetative stem. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – leaves of vegetative stem. Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.Cerastium arvensesubsp.arvense – seeds under magnification. The seeds are less than 1 mm in size, reniformeith tuberculate surface.Osogovo – “Markova stupka” region of subalpine pastures near “Carev vrv” peak, 15.07.2023. Photo by Trajche Dimitrovski.
Jalas, J. (perennial spp.); Sell, P. D. & Whitehead, F. H. (annual spp.). 1964. Cerastium L. in Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A. (eds.), Flora Europaea. Volume 1. Lycopodiaceae to Platanaceae. Cambridge: Cambridge University Press, pp. 136 -145.
Gajić, M. 1970. Rod Cerastium L. In Josifović, M. (ed.), Flora SR Srbije, II. Beograd: Srpska Akademija Nauka i Umetnosti, pp. 156–176.
