Determined according to the botanical keys provided by Micevski in Micevski (1998).
Note: The indumentum of leaves, an important taxonomic trait seems to be variable across the native range and therefore variously described by different authors (Favarger and Zésiger in Tutin et al., 1964; Gajić in Josifović, 1972; Micevski in Micevski, 1998). The populations from Osogovo have persistent indumentum of dense, short glandular hairs on both leaf surfaces and longer and thicker marginal cilia, in accordance with the description provided by Micevski in Micevski (1998).
Distribution: Native to Italy, Balkan Peninsula to Slovakia and Ukraine according to The Euro+Med PlantBase (Marhold, 2011+) and the Plants of the World Online portal (Royal Botanic Gardens, Kew).
Habitat: Rocky terrains – on rocks in the highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region) at c. 1350 m altitude, growing alongside other succulents (Sempervivum erythraeum, Jovibarba heufelii, Sedumstefco, Sedum anuum).
Lifeform: Perennial succulent; Hemicryptophyte according to the Raunkiær classification.
Flowering period: Summer: JUL.
Sempervivum marmoreum – habit. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – habit. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – habit. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – habit. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – habit. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – inflorescence. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – inflorescence. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – inflorescence. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – inflorescence. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – developing inflorescence. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – branch of inflorescence with flowers. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – branch of inflorescence with flowers. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – flower. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – flower. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – flower. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – flower. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – flower. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – flower. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – flower. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – flower and flower buds. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – side view of calyx. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – side view of calyx. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – upper part of stem and branches of cyme. Osogovo- highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stem with cauline leaves. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stem with cauline leaves. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stem with cauline leaves. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stem with cauline leaves. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stem with cauline leaves. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – lower leaves of flowering stem. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – adaxial side of cauline leaf. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – abaxial side of cauline leaf. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stem with glandular hairs. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – indumentum of stem and cauline leaves. a- glandular hairs of stem; b- marginal cilia; c – glands on the abaxial side of leaf; d- glands on the adaxial side of leaf. Osogovo – highest part of the “Ratkova Skala” region bordering with Visoka chuka (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosettes. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosettes. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosettes. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosettes. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosettes. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosettes. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal leaves. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – adaxial side of basal leaves. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – adaxial side of basal leaf. a-glands on the adaxial side; b – marginal cilia. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – abaxial side of basal leaves. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – abaxial side of basal leaf and glands under magnification. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosettes and stolons. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka”(Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosettes and stolons. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka”(Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stolon. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka”(Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stolons. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka”(Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stolon. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka”(Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stolons. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – inflorescence from the previous year with dehisced capsules. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – dehisced capsules. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 08.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – habit. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – habit. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – habit. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – habit. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – inflorescence. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – inflorescence. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – inflorescence. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosettes and stolons. Scale in cm/ mm. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – basal rosette. Scale in cm/ mm. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stolon. Scale in cm/ mm. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – stolons. Scale in cm/ mm. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region), 20.07.2023. Photo by Trajche Dimitrovski.Sempervivum marmoreum – flower size. Scale in cm/ mm. Osogovo – highest part of the “Ratkova Skala” region bordering with “Visoka chuka” (Ponikva region). Photo by Trajche Dimitrovski.
References:
Sempervivum L. Favarger, C. and Zésiger, F. in Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M., & Webb, D.A. (1964). Flora Europaea. Volume 1. Lycopodiaceae to Platanaceae. Cambridge: Cambridge University Press, pp. 352 – 355.
Sempervivum L. Micevski, K. (1998). The Flora of the Republic of Macedonia Vol. I, Book 4. Macedonian Academy of Sciences and Arts, pp. 1042 – 1048.
