Verbascum glabratum subsp. bosnense (K. Malý) Murb.

Taxonomy: Fam. Scrophulariaceae// Genus: Verbascum// Species: Verbascum glabratum Friv.// Subspecies: Verbascum glabratum subsp. bosnense (K. Malý) Murb.

Determined according to the botanical keys provided by Nikolić in Josifović (1974) and Ferguson in Tutin et al. 1972. Differs from other subspecies in having flowers grouped in fascicles of 2 to 7 and basal leaves coarsely crenate or lobed, more or less pinnately incised towards the base.

Lifeform: Perrenial – hemicryptophyte according to the Raunkiær system.

Distribution: Native to the Balkan Peninsula according to the distribution information provided by the the Plants of the World Online portal (Royal Botanic Gardens, Kew).

Habitat: Edges of forests, sparse woods in the oak forests zone.

Flowering period: Spring – summer: MAY, JUN.

Verbascum glabratum subsp. bosnense – habit. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – habit. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – habit. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – habit. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – habit. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – habit. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – plants in vegetative stage. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – plants in vegetative stage. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – inflorescence. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – inflorescence. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – inflorescence. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – branch of inflorescence. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – branch of inflorescence. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – branch of inflorescence. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – flowers. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – flowers. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – flowers. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – flowers. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – flowers. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – side view of flowers and calyx. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – capsules. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – capsules. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – stem of inflorescence, capsules and bract. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – axis of inflorescence, capsules and bract. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – axis of inflorescence. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – stem and upper cauline leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – stem and cauline leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – stem and cauline leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – stem and cauline leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – abaxial side of upper cauline leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – abaxial side of cauline leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – stem. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – basal leaves. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – basal leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – basal leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – adaxial side of basal leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – abaxial side of basal leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – adaxial side of basal leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – petiole of basal leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – petiole of basal leaf. Osogovo – village of Pantelej region, 22.06.2024. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – basal leaves. Scale in cm. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – cauline leaves. Scale in cm. Photo by Trajche Dimitrovski.
Verbascum glabratum subsp. bosnense – A: flower; B: anterior stamens (as) and posterior stamens (ps). All stamens with reniform anthers, not decurrent or obliquely inserted. Filaments with violet hairs, the anterior glabrous in the upper part. C, D and E: fascicles with capsules. br- bract; brl – bracteole. Photo by Trajche Dimitrovski.

References:

Rod Verbascum L. Nikolić, V. in Josifović, M. (1974). Flora SR Srbije VI. Srpska Akademija Nauka i Umetnosti, Beograd, pp. 101 – 142.

Verbascum glabratum subsp. bosnense (K.Malý) Murb. Plants of the World Online, Royal Botanic Gardens, Kew.  https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:77252035-1 [Accessed 24.06.2024].

Verbascum glabratum subsp. bosnense. Marhold, K. (2011+): Verbascum. – In: Euro+Med Plantbase – the information resource for Euro-Mediterranean plant diversity. https://europlusmed.org/cdm_dataportal/taxon/125bfe0e-76e2-48d3-96e6-a4c3660b9926 [Accessed 24.06.2024].

Verbascum L. (incl. Celsia L.). Ferguson, I.K. in Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (1972). Flora Europaea. Volume 3: Diapensiaceae to Myoporaceae. Cambridge: Cambridge University Press, pp. 205 – 216.