Taxonomy: Fam. Dryopteridaceae// Genus: Dryopteris// Species: Dryopteris dilatata (Hoffm.) A. Gray. The classification is given according to The Euro+Med PlanBase (source: Christenhusz & Raab-Straube, 2013).
Determined according to the botanical keys provided by Heywood in Tutin et. al (1964), Heywood revised by Praser-Jenkins in Tutin et al. (1993), Micevski (1985), and Szczęśniak, Tlałka, and Rostański in Szczęśniak and Gola (2009).
Lifeform: Perrenial; Hemicryptophyte according to the Raunkiær system.
Distribution: Native to the Azores, Europe to Iran, Southwest Siberia to Mongolia according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Garden, Kew).
Habitat: Found in moist, humus rich sites along springs in “Jamishko Оsoe” region above “Venechka” River, at c. 1320 m altitude.
Dryopteris dilatata – habit. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – habit. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – habit. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnae. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnae. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnae. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnae. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – rachis and pinnae with pinnules. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – lowest pinnae. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – lowest pinnae. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – acroscopic and basioscopic pinnules of lowest pinnae. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnae with pinnules. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnae with pinnules. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnae with pinnules. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnules. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnules. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – pinnules. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – adaxial side of pinnules. Osogovo – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – adaxial side of pinnules with sori. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – adaxial side of pinnules with sori. Osogovo – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – adaxial side of pinnules with sori. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – adaxial side of pinnules with sori. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – adaxial side of pinnules with sori. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – adaxial side of pinnules with sori. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – adaxial side of pinnules with sori. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata –abaxial side of pinnules and rachis. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata –abaxial side of pinnules and rachis. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – rachis. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – rachis. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – rachis. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – leaf petioles with scales. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata – leaf petioles with scales. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata –leaf petiole with scales. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata –leaf petiole with scales. Osogovo – Ponikva – “Jamisko Osoe” region, 01.10.2025. Photo by Trajche Dimitrovski.Dryopteris dilatata –leaf lamina. The lamina is up to 2 times as long as wide, the proximal basioscopic pinnule of the basal pinnae is less than half as long as the pinna. Specimen from Osogovo – Ponikva – “Jamisko Osoe” region. Scale in cm. Photo by Trajche Dimitrovski.Dryopteris dilatata –leaf. The lamina is up to 2 times as long as wide, the proximal basioscopic pinnule of the basal pinnae is less than half as long as the pinna. Specimen from Osogovo – Ponikva – “Jamisko Osoe” region. Scale in cm. Photo by Trajche Dimitrovski.Dryopteris dilatata –leaf. The lamina is up to 2 times as long as wide, the proximal basioscopic pinnule of the basal pinnae is less than half as long as the pinna. Specimen from Osogovo – Ponikva – “Jamisko Osoe” region. Scale in cm. Photo by Trajche Dimitrovski.Dryopteris dilatata –lowest pinnae. The proximal basioscopic pinnule of the basal pinnae is less than half as long as the pinna, and is often equal or even shorter than the second one. Specimen from Osogovo – Ponikva – “Jamisko Osoe” region. Scale in cm. Photo by Trajche Dimitrovski.Dryopteris dilatata –abaxial side of pinnules with sori. Specimen from Osogovo – Ponikva – “Jamisko Osoe” region. Scale in mm. Photo by Trajche Dimitrovski.Dryopteris dilatata –scales of pedicels. The scales are bicolored, lighter brown at the margin, with a darker brown central stripe up to the apex. Specimen from Osogovo – Ponikva – “Jamisko Osoe” region. Scale in mm. Photo by Trajche Dimitrovski.Dryopteris dilatata –stalked glands of rachis. Photo by Trajche Dimitrovski.
References:
Dryopteris Adanson. Heywood, V.H. in Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M., & Webb, D.A. (1964). Flora Europaea. Volume 1. Lycopodiaceae to Platanaceae. Cambridge: Cambridge University Press, p. 20 – 22.
Dryopteris Adanson. Heywood, V.H. (edition 1) revised by Praser-Jenkins, C.R. in Tutin, T.G., Burges, N.A. Chater, A. O., Edmondson, J. R., Heywood, V. H., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A., assisted by Akeroyd, J.R. and Newton, M.E. (1993). Flora Europaea. Volume 1. Psilotaceae to Platanaceae, second edition. Cambridge: Cambridge University Press, pp. 27 – 30.
Dryopteris Adanson. Micevski, K. (1985). The Flora of SR Macedonia Vol. 1, Book 1. Macedonian Academy of Sciences and Arts, pp. 67 – 72.
Szczęśniak, E., Tlałka, D. and Rostański, A. (2009). Key to identification and descriptions of species of Buckler-ferns (Dryopteris Adans.) occurring in Poland, in Szczęśniak, E. and Gola, E. (eds.) Genus Dryopteris Adans. in Poland. Publisher: Polish Botanical Society & Institute of Plant Biology, University of Wrocław, Wrocław, pp. 5 – 34.
