Determined according to the botanical keys provided by Micevski (1985) and Crabbe et al. in Tutin et al. (1964) as Asplenium onopteris L., Sp. Pl. 1081 (1753). The accepted name in the Euro+Med Plantbase (Christenhusz & Raab-Straube, 2013+) is Asplenium adiantum-nigrum subsp. onopteris (L.) Heufl. in Verh. Zool.-Bot. Vereins Wien 6: 310. 1856.
Lifeform: Perrenial; Hemicryptophyte according to the Raunkiær system.
Distribution: Native to Macaronesia, Ireland to Mediterranean and Iran according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Garden, Kew).
Habitat: Shaded positions – under trees. Recorded in mixed European beech – oak forest at c. 900 m altitude near the waterfall of “Shtalkovicka” River.
Asplenium adiantum-nigrum subsp. onopteris – habit. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – habit. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – habit. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – habit. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – leaf lamina. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – leaf lamina. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – leaf lamina. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – leaf lamina. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – leaf lamina. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – leaf lamina. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – leaf lamina. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – caudate apex of leaf. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – caudate apex of leaf. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – caudate apex of leaf. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – upper pinae. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris –middle pinnae. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – lower pinnae. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – lower pinnae. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – pinnae with caudate apex. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – caudate apex of pinnae. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – caudate apex of pinnae. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – abaxial side of pinnae. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – caudate apex of lowest pinna. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – abaxial side of pinnae and rachis. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – abaxial side of pinnae and rachis. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – abaxial side of pinnae with sori. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – abaxial side of pinnae with sori. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – abaxial side of pinnae with sori. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – petioles. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – petiole. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – petioles. Osogovo – near the waterfall of “Shtalkovichka” River, 02.11.2025. Photo by Trajche Dimitrovski.Asplenium adiantum-nigrum subsp. onopteris – leaf. Specimen from Osogovo – near the waterfall of “Shtalkovichka” River. Scale in cm. Photo by Trajche Dimitrovski.