Trifolium pratense L.

Taxonomy: Fam. Fabaceae// Genus: Trifolium// Species: Trifolium pratense L.

Determined according to the botanical key provided in The Flora of Republic of Macedonia (Micevski, 2001).

Lifeform: Perrenial; Hemicryptophyte according to the Raunkiær system.

Distribution: Native to Northwest Africa, from Europe to Siberia, China (Xinjiang) and India according to the distribution information provided by the Plants of the World Online (Royal Botanic Gardens, Kew).

Habitat:  Grasslands and pastures.

Flowering period:  Late spring – summer: MAY, JUN.

Trifolium pratense – habit. Gorge of  “Kochanska” River, 08.05.2021. Photo by Trajche Dimitrovski.
Trifolium pratense – stem with inflorescence and leaves. Gorge of  “Kochanska” River, 08.05.2021. Photo by Trajche Dimitrovski.
Trifolium pratense – inflorescence and leaf. Gorge of  “Kochanska” River, 08.05.2021. Photo by Trajche Dimitrovski.
Trifolium pratense – stem with inflorescence and leaves. Gorge of  “Kochanska” River, 08.05.2021. Photo by Trajche Dimitrovski.
Trifolium pratense – stem with leaf petiole and stipules. Gorge of  “Kochanska” River, 08.05.2021. Photo by Trajche Dimitrovski.
Trifolium pratense – habit. Gorge of  “Kochanska” River, 17.05.2020. Photo by Trajche Dimitrovski.
Trifolium pratense – stems with inflorescence and leaves. Gorge of  “Kochanska” River, 17.05.2020. Photo by Trajche Dimitrovski.
Trifolium pratense – stem with inflorescence and leaves. Gorge of  “Kochanska” River, 17.05.2020. Photo by Trajche Dimitrovski.

References:

Micevski, K. (2001). The Flora of the Republic of Macedonia Vol. I, Book 5. Macedonian Academy of Sciences and Arts.

Trifolium pratense L. Plants of the World Online, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:523575-1 [Accessed 23.01.2022].

Myosotis ramosissima Rochel

Taxonomy: Fam. Boraginaceae// Genus: Myosotis// Species: Myosotis ramosissima Rochel.

Determined according to the botanical key provided by Matevski and Melovski in Matevski (2010).

Lifeform: Annual; Therophyte according to the Raunkiær system.

Distribution: Native to Macaronesia, Northwest Africa, Europe to Iraq according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Gardens, Kew).

Habitat: Grasslands, pastures, bushlands, river gorges, rocky terrains.

Flowering period: Spring: MAR, APR, MAY.

Myosotis ramosissima – habit. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – habit. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – habit. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – habit. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – inflorescence. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – inflorescence with flowers. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – flowers. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – flowers. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – flowers. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – fruiting calyces. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – fruiting calyces. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – fruiting calyces. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – stem with cauline leaves. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – stem with cauline leaves. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – stem with cauline leaves. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – stem with cauline leaves. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – lower part of plant. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – lower cauline leaves and basal leaves. Hills near Kochani, between the gorges of “Kochanska” River and “Orizarska” River, 16.04.2022. Photo by Trajche Dimitrovski.
Myosotis ramosissima – habit. Gorge of “Kochanska” River,25.04.2021. Photo by Trajche Dimitrovski.
Myosotis ramosissima – habit. Gorge of “Kochanska” River, 06.04.2021. Photo by Trajche Dimitrovski.
Myosotis ramosissima – habit. Gorge of “Kochanska” River, 06.04.2021. Photo by Trajche Dimitrovski.
Myosotis ramosissima – inflorescence. Gorge of “Kochanska” River, 06.04.2021. Photo by Trajche Dimitrovski.
Myosotis ramosissima – inflorescence. Gorge of “Kochanska” River,25.04.2021. Photo by Trajche Dimitrovski.
Myosotis ramosissima – inflorescence. Gorge of “Kochanska” River,25.04.2021. Photo by Trajche Dimitrovski.
Myosotis ramosissima – inflorescence. Gorge of “Kochanska” River,25.04.2021. Photo by Trajche Dimitrovski.
Myosotis ramosissima – inflorescence. Gorge of “Kochanska” River, 06.04.2021. Photo by Trajche Dimitrovski.
Myosotis ramosissima – stem with cauline leaves. Gorge of “Kochanska” River,25.04.2021. Photo by Trajche Dimitrovski.
Myosotis ramosissima – lower part of stem with cauline leaves. Gorge of “Kochanska” River, 06.04.2021. Photo by Trajche Dimitrovski.

References:

Myosotis ramosissima Rochel. Plants of the World Online, Royal Botanic Gardens Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:119209-1 [Accessed 21.01.2022].

Myosotis L. Matevski, V. and Melovski, Lj. in Matevski (2010). The Flora of the Republic of Macedonia Vol. II, Book 1. Macedonian Academy of Sciences and Arts, pp. 141 – 163.

Last update: 29.01.2023.

Parentucellia latifolia (L.) Caruel

Taxonomy: Fam. Orobanchaceae// Genus: Parentucellia// Species: Parentucellia latifolia (L.) Caruel. The classification is given according to the Euro+Med Planbase (Marhold, 2011+).

Determined according to the botanical keys provided in Flora Europaea Vol. 3 (Tutin et al., 1972).

Lifeform: Hemiparasitic annual plant; therophyte according to the Raunkiær system.

Distribution: Native to Canary Islands, Mediterranean to Afghanistan, Sudan (Jebel Marra) according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Gardens, Kew).

Habitat:  Open habitats- grasslands/ pastures, rocks and rocky terrains, bushland, alongside roads.

Flowering period:  Spring: APR, MAY.

Parentucellia latifolia – habit. Gorge of “Kochanska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Parentucellia latifolia – habit. Gorge of “Kochanska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Parentucellia latifolia – habit. Gorge of “Kochanska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Parentucellia latifolia – stem with inflorescence and cauline leaves. Gorge of “Kochanska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Parentucellia latifolia – stem with inflorescence and cauline leaves. Gorge of “Kochanska” River, 25.04.2021. Photo By Trajche Dimitrovski.
Parentucellia latifolia – stem with inflorescence and cauline leaves. Gorge of “Kochanska” River, 25.04.2021. Photo By Trajche Dimitrovski.
Parentucellia latifolia – stem with inflorescence and cauline leaves. Gorge of “Kochanska” River, 25.04.2021. Photo By Trajche Dimitrovski.
Parentucellia latifolia – basal and lower caline leaves. Gorge of “Kochanska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Parentucellia latifolia – habit. Osogovo, grassland near “Saint Panteleimon” monastery, 23.05.2021. Photo by Trajche Dimitrovski.
Parentucellia latifolia – habit. Osogovo, grassland near “Saint Panteleimon” monastery, 23.05.2021. Photo by Trajche Dimitrovski.
Parentucellia latifolia – inflorescence with fruits and flowers. Osogovo, grassland near “Saint Panteleimon” monastery, 23.05.2021. Photo by Trajche Dimitrovski.
Parentucellia latifolia – inflorescence. Osogovo, grassland near “Saint Panteleimon” monastery, 23.05.2021. Photo by Trajche Dimitrovski.
Parentucellia latifolia – inflorescence. Osogovo, grassland near “Saint Panteleimon” monastery, 23.05.2021. Photo by Trajche Dimitrovski.
Parentucellia latifolia – stem with cauline leaves. Osogovo, grassland near “Saint Panteleimon” monastery, 23.05.2021. Photo by Trajche Dimitrovski.
Parentucellia latifolia – habit (right) and and inflorescence (left). Gorge of “Kochanska” River, 05/ 2020. Photo By Trajche Dimitrovski.
Parentucellia latifolia – flowers. Gorge of “Kochanska” River, 05/ 2020. Photo By Trajche Dimitrovski.
Parentucellia latifolia – flowers. Gorge of “Kochanska” River, 05/ 2020. Photo By Trajche Dimitrovski.

References:

Parentucellia latifolia (L.) Caruel. Plants of the World Online, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:806703-1 [Accessed 20.01.2022].

Parentucellia latifolia. Marhold, K. (2011+): Parentucellia. – In: Euro+Med Plantbase – the information resource for Euro-Mediterranean plant diversity. Published on the Internet http://ww2.bgbm.org/EuroPlusMed/PTaxonDetail.asp?NameId=34247&PTRefFk=7200000 [Accessed 20.01.2022].

Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (1972). Flora Europaea. Volume 3: Diapensiaceae to Myoporaceae. Cambridge: Cambridge University Press.

Minuartia hirsuta subsp. falcata (Griseb.) Mattf.

Taxonomy: Fam. Caryophyllaceae// Genus: Minuartia// Species: Minuartia hirsuta (M. Bieb.) Hand.-Mazz.// Subspecies: Minuartia hirsuta subsp. falcata (Griseb.) Mattf.

Determined according to the botanical key provided in The Flora of the Republic of Macedonia (Micevski, 1993).

Distribution: Minuartia hirsuta (M. Bieb.) Hand.-Mazz. is native from Central Europe to Iran according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Gardens Kew).

Habitat: Rocks and rocky terrains.

Lifeform: Perennial; Hemicryptophyte according to the Raunkiær system.

Flowering period: Spring-summer: APR, MAY, JUN, JUL.

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Minuartia hirsuta – habit. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – habit. Gorge of “Kochanska” River, 25.04.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – habit. Gorge of “Kochanska” River, 25.04.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – flowers. Gorge of “Kochanska” River, 25.04.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – habit. Gorge of “Kochanska” River, 20.04.2021. Photo By Trajche Dimitrovski.
Minuartia hirsuta – habit. Gorge of “Kochanska” River, 20.04.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – inflorescence. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – inflorescence. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – flowers. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – flowers. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – flowers. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – inflorescence. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
Minuartia hirsuta – flowers. Gorge of “Kochanska” River, 25.04.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – flowers. Gorge of “Kochanska” River, 25.04.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – flowers. Gorge of “Kochanska” River, 20.04.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – flowers. Gorge of “Kochanska” River, 20.04.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – flowering stems with developing inflorescence and cauline leavea. Gorge of “Kochanska” River, 20.04.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – stems with leaves. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – vegetative stems. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – plant base with vegetative stems. Gorge of “Kochanska” River, 20.06.2021. Photo By Trajche Dimitrovski.
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Minuartia hirsuta – plant base and stems with leaves. Gorge of “Kochanska” River, 20.04.2021. Photo By Trajche Dimitrovski.
Minuartia hirsuta – habit. Gorge of “Kochanska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Minuartia hirsuta – habit. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Minuartia hirsuta – flower. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Minuartia hirsuta – flower. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Minuartia hirsuta – calyx of flower. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 11.04.2021. Photo By Trajche Dimitrovski.
Minuartia hirsuta – stems with leaves. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 11.04.2021. Photo By Trajche Dimitrovski.

References:

Micevski, K. (1993). The Flora of the Republic of Macedonia Vol. 1, Book 2. Macedonian Academy of Sciences and Arts.

Minuartia hirsuta (M.Bieb.) Hand.-Mazz. Plants of the World Online portal, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:155537-1 [Accessed 19.01.2022].

Erodium cicutarium (L.) L’Hér.

Taxonomy: Fam. Geraniaceae// Genus: Erodium// Species: Erodium cicutarium (L.) L’Hér.

Determined according to the botanical keys provided by Webb and Chater in Tutin et al. (1968) and Micevski in Micevski (2005).

Lifeform: Annual; Therophyte according to the Raunkiær system.

Distribution: Native to Macaronesia, Temperate Eurasia to North and Northeast Africa according to the Plants of the World Online portal.

Habitat:  Grasslands/ pastures, rocky terrains, suburban areas, near roads.

Flowering period:  Spring-summer: MAR, APR, MAY, JUN, JUL.

Erodium cicutarium – habit. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – habit. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – habit. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – stem and inflorescence. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – inflorescence. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – inflorescence. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – flower. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – flower. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – flowers. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – flowers. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – calyx of flower. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – fruits. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – fruits. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – fruits. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – peduncle. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – stem with leaves. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – adaxial side of leaf. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – leaf. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – leaf. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – lower part of stem. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – adaxial side of leaf. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – adaxial side of leaf. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – adaxial side of leaf. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – abaxial side of leaf. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – abaxial side of leaf. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – abaxial side of leaf. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – stem. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium – stem. Osogovo – gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Erodium cicutarium– habit. Osogovo, hilly region near village of Beli, 24.05.2021. Photo by Trajche Dimitrovski.
Erodium cicutarium– habit in habitat. Osogovo, hilly region near village of Beli, 06.03.2021. Photo by Trajche Dimitrovski.
Erodium cicutarium– habit. Osogovo, hilly region near village of Beli, 06.03.2021. Photo by Trajche Dimitrovski.
Erodium cicutarium– habit. Osogovo, hilly region near village of Beli, 06.03.2021. Photo by Trajche Dimitrovski.
Erodium cicutarium– inflorescence with flowers. Osogovo, hilly region near village of Beli, 06.03.2021. Photo by Trajche Dimitrovski.
Erodium cicutarium– flower. Osogovo, hilly region near village of Beli, 06.03.2021. Photo by Trajche Dimitrovski.
Erodium cicutarium– fruits. Osogovo, hilly region near village of Beli, 06.03.2021. Photo by Trajche Dimitrovski.
Erodium cicutarium– fruits. Osogovo, hilly region near village of Beli, 06.03.2021. Photo by Trajche Dimitrovski.
Erodium cicutarium– plant base with leaves. Osogovo, hilly region near village of Beli, 06.03.2021. Photo by Trajche Dimitrovski.
Erodium cicutarium– leaves. Osogovo, hilly region near village of Beli, 06.03.2021. Photo by Trajche Dimitrovski.

References:

Erodium ciconium (L.) L’Hér. Plants of the World Online portal, Royal Botanic Gardens,Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:372219-1 [Accessed 19.01.2022].

Erodium L’Hér. Micevski, K. in Micevski, K. (2005). The Flora of the Republic of Macedonia Vol. I, Book 6. Macedonian Academy of Sciences and Arts, pp. 1506 – 1517.

Erodium L’Hér. Webb, D.A. and Chater, A.O. in Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (1968). Flora Europaea. Volume 2: Rosaceae to Umbelliferae. Cambridge: Cambridge University Press, pp. 199 -204.

Last update: 06.12.2023.

Silene latifolia Poir. (syn. Silene alba (Miller) Krause)

Taxonomy: Fam. Caryophyllaceae// Genus: Silene// Species: Silene latifolia Poir.

Determined according to the botanical key provided in The Flora of the Republic of Macedonia (Micevski, 1993) as Silene alba (Miller) Krause, in Sturm, Deutschl. Fl. ed. 2 (5): 98 (1901). The accepted name in The Euro+Med PlantBase (source: Marhold, 2011) is Silene latifolia Poir., Voy. Barbarie 2: 165. 1789.

Distribution: Native to Temperate Eurasia, Northwest Africa according to the distribution information provided by the Plants of the World Online, Royal Botanic Gardens, Key.

Lifeform: Perrenial; hemicryptophyte according to the Raunkiær system.

Habitatat: Edges and clearings of forests, woods, alongside roads.

Silene latifolia – habit. Gorge of “Kochanska” River, 08.05.2021. Photo by Trajche Dimitrovski.

Flowering period: Spring-summer: APR, MAY, JUN, JUL, AUG.

Silene latifolia – habit in habitat. Gorge of “Kochanska” River, 03.04.2021. Photo by Trajche Dimitrovski.
Silene latifolia – habit. Gorge of “Kochanska” River, 08.05.2021. Photo by Trajche Dimitrovski.
Silene latifolia – male inflorescence. Gorge of “Kochanska” River, 03.04.2021. Photo by Trajche Dimitrovski.
Silene latifolia – male flower. Gorge of “Kochanska” River, 03.04.2021. Photo by Trajche Dimitrovski.
Silene latifolia – male flower. Gorge of “Kochanska” River, 03.04.2021. Photo by Trajche Dimitrovski.
Silene latifolia – male flower. Gorge of “Kochanska” River, 08.05.2021. Photo by Trajche Dimitrovski.
Silene latifolia – male flower. Gorge of “Kochanska” River, 08.05.2021. Photo by Trajche Dimitrovski.
Silene latifolia – female flower. Gorge of “Kochanska” River, 03.04.2021. Photo by Trajche Dimitrovski.
Silene latifolia – female flower. Gorge of “Kochanska” River, 03.04.2021. Photo by Trajche Dimitrovski.
Silene latifolia female plant- habit (left) and dichasium (right). Osogovo, lake “Gradche”, 12.05.2020. Photo by Trajche Dimitrovski.
Silene latifolia – female flowers. Osogovo, lake “Gradche”, 12.05.2020. Photo by Trajche Dimitrovski.
Silene latifolia – habit. Gorge of “Kochanska” River, 28.05.2020. Photo by Trajche Dimitrovski.
Silene latifolia – male flowers. Gorge of “Kochanska” River, 28.05.2020. Photo by Trajche Dimitrovski.

References:

Micevski, K. (1993). The Flora of the Republic of Macedonia Vol. I, Book 2. Macedonian Academy of Sciences and Arts.

Silene latifolia. Marhold, K. (2011): Caryophyllaceae. – In: Euro+Med Plantbase – the information resource for Euro-Mediterranean plant diversity. Published on the Internet http://ww2.bgbm.org/EuroPlusMed/PTaxonDetail.asp?NameId=102239&PTRefFk=7200000 [Accessed 19.01.2022].

Silene latifolia Poir. Plants of the World Online, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:157647-1 [Accessed 19.01.2022].

Alkanna tinctoria Tausch

Taxonomy: Fam. Boraginaceae// Genus: Alkanna// Species: Alkanna tinctoria Tausch.

Determined according to the botanical key provided in The Flora of the Republic of Macedonia (Matevski, 2010).

Distribution: Native from the Mediterranean to South Central Europe according to the distribution information provided by the Plants of the World Online portal.

Lifeform:  Perennial; Hemicryptophyte according to the Raunkiær system.

Habitat: Dry and open habitats- grasslands, pastures and rocky terrains.

Flowering period:  Spring: MAR, APR, MAY.

Alkanna tinctoria – habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – habit. Osogovo, mountain ridge between the gorge of “Kochanska” River and “Orizarska” River, 10.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – habit. Osogovo, mountain ridge between the gorge of “Kochanska” River and “Orizarska” River, 10.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – habit. Osogovo, mountain ridge between the gorge of “Kochanska” River and “Orizarska” River, 10.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – flowers. Osogovo, mountain ridge between the gorge of “Kochanska” River and “Orizarska” River, 10.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – flowering stem with inflorescence. Osogovo, mountain ridge between the gorge of “Kochanska” River and “Orizarska” River, 10.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – flowers. Osogovo, mountain ridge between the gorge of “Kochanska” River and “Orizarska” River, 10.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – habit in habitat. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – habit. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – habit. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – habit. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – flowering stem with inflorescence. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – flowering stem with inflorescence. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – flowering stem with inflorescence. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – inflorescence. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – inflorescence. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria – inflorescence. Hills on the periphery of Kochani, 06.04.2021. Photo by Trajche Dimitrovski.   
Alkanna tinctoria (habit). Osogovo Mountains, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Alkanna tinctoria (habit). Osogovo Mountains, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Alkanna tinctoria – inflorescence. Osogovo, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Alkanna tinctoria – inflorescence. Osogovo, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Alkanna tinctoria – inflorescence. Osogovo, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Alkanna tinctoria – flowers. Osogovo, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Alkanna tinctoria – stem with cauline leaves. Osogovo Mountains, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Alkanna tinctoria (basal rosette). Osogovo Mountains, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Alkanna tinctoria – basal leaves. Osogovo, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Alkanna tinctoria – basal leaves. Osogovo, “Kochanski dol” near Kochani, 05.07.2020. Photo by Trajche Dimitrovski

References:

Alkanna tinctoria (L.) Tausch. Plants of the World Online, Royal Botanic Gardens Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:112753-1 [Accessed 19.01.2022].

Matevski, V. (2010). The Flora of the Republic of Macedonia Vol. II, Book 1. Macedonian Academy of Sciences and Arts.

Viola reichenbachiana Boreau

Taxonomy: Fam. Violaceae// Genus: Viola// Species: Viola reichenbachiana Boreau

Determined according to the botanical key provided by of Macedonia (Micevski, 1995).

Lifeform: Perrenial; Hemicryptophyte according to the Raunkiær system.

Distribution: Native from Europe to Iran, Northwest Africa according to the distribution information provided by the Plants of the World Online portal.

Habitat:  Edges of forests, woods.

Flowering period: Spring: APR,MAY.

Viola reichenbachiana – habit. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower with calyx and spur. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower with calyx and spur. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower with calyx and spur. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower with calyx and spur. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower bud. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flowering stems stems with cauline leaves and basal leaves at plant base. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flowering stems stems with cauline leaves and basal leaves at plant base. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flowering stems with cauline leaves. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – cauline leaves. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – stems and stipules. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – stems and stipules. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – lower part of stem and basal leaves. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – lower part of stem and basal leaves. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – plant base. “Shtalkovichka” River near village of “Shtalkovica”, 25.04.2022. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flowers. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – cauline leaves and stipules. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – stem and stipules. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – basal leaves. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – basal leaves. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – adaxial side of leaf. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – abaxial side of leaf. Gorge of “Kochanska” River, 02.04.2023. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – habit. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – flower. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – capsule. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.
Viola reichenbachiana – leaf. Gorge of “Kochanska” River, 26.04.2021. Photo by Trajche Dimitrovski.

References:

Viola L. Micevski, K. (1995). The Flora of the Republic of Macedonia Vol. I, Book 3. Macedonian Academy of Sciences and Arts, pp. 503- 547.

Viola reichenbachiana Boreau. Plants of the World Online portal, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:869090-1 [Accessed 18.01.2022].

Last update: 09.04.2023.

Viola kitaibeliana Schult.

Taxonomy: Fam. Violaceae// Genus: Viola// Species: Viola kitaibeliana Schult.

Determined according to the botanical key provided in The Flora of the Republic of Macedonia (Micevski, 1995).

Lifeform: Annual; therophyte according to the Raunkiær system.

Distribution: Native to Canary Islands, France, South Europe, Central Europe to Iran and Turkmenistan, Northwest Africa according to the distribution information provided by the Plants of the World Online portal.

Habitat:  Grasslands/ pastures, rocky terrains, river gorges, disturbed ground.

Flowering period: Spring: MAR, APR,MAY.

Viola kitaibeliana – habit. Kochani, 02.04.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit. Kochani, 02.04.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit. Kochani, 19.03.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit. Kochani, 19.03.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit. Kochani, 19.03.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit. Kochani, 19.03.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – flower. Kochani, 19.03.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit in habitat. Gorge of ‘Kochanska” River, 27.03.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit. Gorge of ‘Kochanska” River, 27.03.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit in habitat. Gorge of ‘Kochanska” River, 01.04.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit in habitat. Gorge of ‘Kochanska” River, 01.04.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit. Gorge of ‘Kochanska” River, 01.04.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit. Gorge of ‘Kochanska” River, 01.04.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – habit at the start of flowering. Gorge of ‘Kochanska” River, 13.03.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – flower. Gorge of ‘Kochanska” River, 01.04.2021. Photo taken by Trajche Dimitrovski.
Viola kitaibeliana – flower. Gorge of ‘Kochanska” River, 13.03.2021. Photo taken by Trajche Dimitrovski.

References:

Micevski, K. (1995). The Flora of the Republic of Macedonia Vol. I, Book 3. Macedonian Academy of Sciences and Arts.

Viola kitaibeliana Schult. Plants of the World Online portal, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:868446-1 [Accessed 18.01.2022].

Echinops microcephalus Sm.

Taxonomy: Fam. Compositae// Genus: Echinops// Species: Echinops microcephalus Sm.

Determined according to the botanical key provided by Kožuharov in Tutin et al. (1976).

Lifeform: Perrenial; hemicryptophyte according to the Raunkiær system.

Distribution: Native from Southeast Europe to Turkey according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Gardens, Kew).

Habitat:  Dry and open habitats – dry grassland, clearings of oak forests, hill pastures.

Flowering period: Summer – fall: AUG, SEP.

Echinops microcephalus – habit. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – habit. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – habit. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – habit. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – inflorescence with capitula. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – inflorescence with capitula. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – inflorescence with capitula. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – inflorescence with capitula. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – inflorescence with capitula. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – inflorescence with capitula. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – spent capitula. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – stem with leaves. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – stem with leaves. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – stem with leaves. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – leaves. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – leaves. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – leaf. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – adaxial side of leaf. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – adaxial side of leaf. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – stem. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – lower part of stems. Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – inflorescence. Scale in cm/mm. Specimen from Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – capitula. Scale in mm. Specimen from Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – capitula. Scale in mm. Specimen from Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – floret (left) and involucral bracts (right). Scale in mm. Specimen from Osogovo – near village of Nivichani, 13.08.2023. Photo by Trajche Dimitrovski.
Echinops microcephalus – habit. Osogovo, hilly region near village of Beli, 12.09.2021. Photo by Trajche Dimitrovski.
Echinops microcephalus – habit. Osogovo, hilly region near village of Beli, 12.09.2021. Photo by Trajche Dimitrovski.
Echinops microcephalus – inflorescence with capitula. Osogovo, hilly region near village of Beli, 12.09.2021. Photo by Trajche Dimitrovski.
Echinops microcephalus – inflorescence with spent capitula. Osogovo, hilly region near village of Beli, 12.09.2021. Photo by Trajche Dimitrovski.
Echinops microcephalus – stem with cauline leaves. Osogovo, hilly region near village of Beli, 12.09.2021. Photo by Trajche Dimitrovski.
Echinops microcephalus – cauline leaves. Osogovo, hilly region near village of Beli, 12.09.2021. Photo by Trajche Dimitrovski.

References:

Echinops microcephalus Sm. Plants of the World Online portal, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:202585-1 [Accessed 17.01.2022].

Echinops L. Kožuharov, S. in Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (1976). Flora Europaea. Volume 4: Plantaginaceae to Compositae (and Rubiaceae). Cambridge: Cambridge University Press, pp. 212 – 214.

Last update: 16.02.2024.